Ruff - male displaying on lek (Philomachus pugnax). Sweden. Extremely sexually dimorphic. Breeds in Arctic and subarctic and south to termperate zones in Europe. Requires habitat with adjacent feeding-lekking and nesting areas, coastal tundra to forest tundra, near small lakes and in marshes. Migratory with chief winter quarters in Africa. Eats terrestrial and aquatic insects.